The springs push back with f(Δ) = Δ + βΔ³ when stretched by Δ, so each one stiffens as it stretches: its stiffness is 1 + 3βΔ² (line thickness). That is what couples the modes, exactly as stretching the midplane couples the modes of a plate. The strike gives the marked masses a sudden velocity; the sound is the velocity of the mass at the pickup ring.